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Her tricky smile and timeless allure have inspired academic study and artistic emulation for more than five centuries. But the story of this perplexing portrait of 'Mona Lisa' is even richer than it looks.

United statesFrance generalPablo picassoUngaza villegasMargaret livingstoneMona lisaLady lisaMonna lisaLisa gherardiniVincenzo peruggiaKarl deckerDianne halesAssociated pressUniversity of freiburgFrancesco del giocondoTuileries palace