2The Center for Genome Architecture, Baylor College of Medicine, Houston, TX 77030, USA. 3Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, TX 77030, USA. 4Center for Theoretical Biological Physics, Rice University, Houston, TX 77005, USA. 5Division of Cell Biology, Oncode Institute, Netherlands Cancer Institute, 1066 CX Amsterdam, Netherlands. 6Division of Gene Regulation, Oncode Institute, Netherlands Cancer Institute, 1066 CX Amsterdam, Netherlands. 7Department of Physics, Institute of Biosciences, Letters and Exact Sciences, São Paulo State University (UNESP), São José do Rio Preto – SP, 15054-000, Brazil. 8BioImaging Facility, Netherlands Cancer Institute, 1066 CX Amsterdam, Netherlands. 9Shanghai Institute for Advanced Immunochemical Studies, ShanghaiTech, Pudong 201210, China.