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Billions of years ago, self-replicating systems of molecules became separated from one another by membranes, resulting in the first cells. Over time, evolving cells enriched the living world with an astonishing diversity of new shapes and biochemical innovations, all made possible by compartments. Compartmentalization is how all living systems are organized today from proteins and small molecules sharing space in separate phases to dividing labor and specialized functions within and among cells. ....